Abundance: 0.01 to 82.20% (median 3.27%)
Latitudinal range: -37.0° to 51.9°Habitats: (44) only
Found in 44 samples
Australia: Grant Hall (Layer 2), Grant Hall (Layer 3)
Brazil: Lapa do Santo (Early Holocene), Lapa do Santo (Middle Holocene)
Ecuador: La Chimba, Challuahamba
Ethiopia: Bodo D'ar
France: Saint-Privat-des-Vieux
Greece: Lakonis Cave I (hearth complex), Lakonis Cave I (upper bone breccia), Lakonis Cave I (lower bone breccia)
India: Chandankhera (Period III: Satavahana), Chandankhera (Period IV: Vakataka)
Italy: Rifreddo, La Sassa Cave, Pirro Nord 13
Kazakhstan: Tasbas (Phase 2a)
Kenya: Usenge 3 (Urewe)
Mexico: El Zurdo
Romania: Noviodunum (TPF sector, Middle Ages)
Russia: Kuznetsovka
South Africa: Plovers Lake
Spain: Fuente Nueva 3 (layer 5), Venta Micena 4-I, Venta Micena 4-II, Barranco Léon
Tanzania: Ewass Oldupa (Trench 2), Ewass Oldupa (Trench 3), Ewass Oldupa (Trench 5)
Thailand: Ban Rai rockshelter, Tham Lod rockshelter (Unit 1), Tham Lod rockshelter (Unit 2), Tham Lod rockshelter (Unit 3)
United States: Vess Cave, McKinley Site (Zone A), McKinley Site (Zone B), Steck, Deadman Slough (Excavation Unit), Sucices Site (Excavation Square 2), Watson Brake (3.2 mm fraction), Rattlesnake Point Pueblo (Unit 2), Hale Farm, Pine Hill, Kincaid Shelter
See also Testudines indet.